adephaga aedeagus aerotrophic appendiculate basal caudal clypeus clathrate coarctate connate coxae cryptogastrous cryptonephric deflexed emarginate emargination empodium epipleura eruciform explanate flabellate frons galea glabrous gular heteromorphosis hypsotactic isomerous labrum lacinia lanceolate malpighian maxillae mentum mesepimera mesepisterna metasterna metasternum mola moniliform myrmecophile notopleural oblongum onisciform ovarioles pectinate pedicel pedunculate pinctate planidium prognathous pronotum prosternal pseudotetramerous pubescent pygidium quadrate rostrate scape scrobe securiform striae striole subcylindrical submentum submoniliform subvertical trochanter trochantins urogomphi